@Cassie_Raby These were later followed up and developed by Duboscq et al. (2016) for the louse estimation.
Here are my references:
Tanaka & Takefushi (1993). https://t.co/Ax02joqgZU
Zamma (2002). https://t.co/TpaqMZv3n8
Duboscq (2016). https://t.co/6nVgf1prRr and https://t.co/ZeRiNAvQaf