著者

HORI KIYOTAKA EBIHARA ATSUSHI NAKATO NARUMI MURAKAMI NORIAKI

出版者

日本植物分類学会

雑誌

APG : Acta phytotaxonomica et geobotanica (ISSN:13467565)

巻号頁・発行日

vol.66, no.1, pp.47-57, 2015-02-27

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The Dryopteris varia complex (subg. Erythrovariae, sect. Variae Fraser-Jenk.) is an apogamous fern complex. Most species of the complex are apogamous triploids; Dryopteris saxifraga and D. varia, the only known diploid sexual cytotypes in the complex, are insufficient to account for the morphological and genetic variation within the complex. In this study, we describe a new diploid sexual species, Dryopteris protobissetiana K. Hori et N. Murak., from southern Yakushima, and discuss its morphological characteristics, reproductive mode, and chromosome number. Dryopteris protobissetiana is similar to D. bissetiana in having slightly bullate scales and a dark green lamina surface, but it differs in having flat, serrate margins at the apex of the upper pinnae. Dryopteris protobissetiana is a sexual diploid with 2n=82, and may be a source of the genetic variation in the apogamous complex.